In population composition,heliophytes increased and shade-tolerant decreased with forest fragmentation. 种群的变化趋势是阴湿或喜阴湿的蕨种的种群趋于减少,而林缘和林窗阳性或喜阳性的蕨种的种群趋于增大。
Betula ermanii and Alnus mandshurica predominated in the colonizing woody species on forest boundaries. 林缘入侵物种主要由岳桦和东北赤杨组成。