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- Achenes ellipsoid-reniform, markedly rugose. 腎形瘦果橢圓體，顯著具皺紋。
- Achenes yellow-brown, ovoid, markedly rugose. 瘦果黃褐色，卵圓形，顯著具皺紋。
- Seeds curved, rugose, endosperm absent. 種子弄彎，具皺紋，無胚乳。
- Seeds nitid, sometimes somewhat rugose or rough. 有光澤的種子，有時有點具皺紋或的粗糙的。
- Seed: testa transversely rugose or glandular punctate. 種子：種皮橫向具皺紋或者腺具點的。
- Achenes subreniform, ca. 1 mm in diam., inconspicuously rugose. 近腎形的瘦果，直徑約1毫米，不明顯具皺紋。
- Seeds convex on 1 face, transversely rugose, with fleshy funicle. 種子凸的在1邊，橫向具皺紋，具肉質的珠柄。
- Leaves petiolate;leaf blade simple, crenate, often rugose. 葉具葉柄單葉，具圓齒，通常具皺紋。
- Seeds few, brown, oblong, compressed, 3-5 mm, slightly rugose. 種子很少，棕色，長圓形，壓扁，3-5毫米，稍具皺紋。
- Seeds numerous;seed coat rugose and scrobiculate-reticulate. 種子多數種皮具皺紋和具小網眼網狀。
- Seeds deltoid-obconic, ca. 3 mm, densely transversely rugose. 種子正三角形倒圓錐形，約3毫米，濃密橫向具皺紋。
- Fruit a capsule, obtusely trigonous, transversely rugose, loculicidal. 果一蒴果，鈍三棱，橫向具皺紋，室背開裂。
- Utricles brownish black, obovate, 1.2-1.5 mm, rugose, circumscissile. 胞果棕黑色，倒卵形的，1.;2-1
- Branches grayish brown, terete, rugose, ferruginous pilosulose when young. 分枝淡灰棕色，圓柱狀，具皺紋，幼時鐵鏽色具小柔毛。
- Leaf blade adaxially deeply rugose, margin revolute, veins deeply impressed. 深深的深深的葉片正面具皺紋，邊緣外卷，脈凹陷。
- Capsule rugose, without wings, echinate or lobed, opening by 4 or 5 lobes at maturity. 蒴果具皺紋，沒有翅，具刺或淺裂的，以成熟時4或5裂片打開。
- Fruit rugose, without prickles and wings, not globose, opening by 4 or 5 lobes at maturity. 果具皺紋，沒有皮刺和翅，不球狀，成熟時以4或5裂片打開。
- Seeds 2, black, ovoid, ca. 0.5 mm in diam., slightly rugose, without strophiole. 種子2，黑色，卵球形，直徑約0.;5毫米，稍具皺紋，沒有種阜。
- Branchlets grayish white when dry, rugose, glabrous, subglabrous, or densely white-hairy. 小枝幹燥時淺灰色白色的，具皺紋，無毛，近無毛，或者濃密白色有毛。
- Leaf blade veins adaxially deeply impressed, surface rugose; bracteoles and sepals 12-17, inside velutinous. 正面深深凹陷的葉片脈，表面具皺紋；小苞片和萼片12-17，裡面短絨毛。
