In population composition,heliophytes increased and shade-tolerant decreased with forest fragmentation. 種群的變化趨勢是陰濕或喜陰濕的蕨種的種群趨於減少,而林緣和林窗陽性或喜陽性的蕨種的種群趨於增大。
Betula ermanii and Alnus mandshurica predominated in the colonizing woody species on forest boundaries. 林緣入侵物種主要由岳樺和東北赤楊組成。