Utricle beak above middle divided into 2 small, acicular beaks, each with a proximal, lateral, recurved or inflexed, firm, spinelike, glabrous process.

• 胞果具喙在中部以上全裂成为小的，针状的喙，每个具一下部的，侧生的，内弯或内折的，坚固的、刺状的无毛的突起。(2