To their surprise, Woolsey and Van der Loos found that cortical layer IV contained multiple distinct clusters of CO-rich neurons in a well-delineated arrangement of rows and columns.

• 染色的结果让伍尔西与凡德路斯相当惊讶，他们发现在第四层皮质中，有许多富含CO的神经元形成群集，并呈现行列分明的阵状排列。