The system could move forwards at the speed of 1 m/a or the patches shifted their sites each other, which led to the clone population and maintained its stable state in even large area.

• 同时，林缘扩散种群具有密度大、生产力高等特征，但其生物量积累和自然稀疏过程与有性植物种群具有相同的规律。