The pedicle retained coelomic cavity and not completely filled with a connective tissue as proposed by Popov, L.E (1992).

• 早期有铰类肉茎腔的存在表明早期有铰类肉茎与无铰类肉茎可能同源，支持了有铰类和无铰类腕足动物的单系起源。