The environmental factors contributed to the differentiation of wild rice into two subspecies are thought to be the water and temperature. Regimes and the role of Rayada, glabrous and Aus types were briefly discussed.

• 演化的环境因素可能是水分与温度,文中就光壳稻和Rayada的酶谱构成型和Aus典型在前文聚类中的地位进行了初步讨论。