The borderlines of the different biome ecotones and the gradients of species distribution in the biome ecotone have been investigated through the method of the quantitative ecology.

• 在大尺度水平上，边缘效应测度的研究主要是应用数量生态学等方法，研究不同气候带之间界线的划分及其物种分布的梯度规律性。