Spikelet fertility was controlled by embryosac fertility, pollen fertility and anther dehiscence. The latter two determined the number of germinating pollen on stigma and affected indirectly the fertilization of spikelets.

• 杂种的小穗育性主要受胚囊育性、粉育性及花药开裂特性所控制，后两者通过影响柱头上花粉的萌发量而决定小穗的受精。