In the Fillicales the leptosporangium has a specialized spore dispersal mechanism involving an annulus of cells with thick walls that ejects s pores explosively as it dries out.

• 蕨类植物中薄囊蕨有一个专门的孢子传播装置，在孢子囊壁的部分细胞壁上不均匀加厚成环带，当孢子成熟后将其散射出。