In addition to describing their morphology, he speculated on a role for glia in sleep and wakefulness and even in executive brain functions such as attention.

• 除描述它们的形态之外，他还推测神经胶质在睡眠和觉醒中有一定作用，甚至推测它们在执行大脑功能比如注意力时也有作用。