He suggested that bees which make hexagonal cells evolved in steps from bees that made round cells, under pressure from natural selection to economise wax.

• 他认为，建筑六角形蜂房的蜜蜂是从建筑圆形蜂房的蜜蜂进化而来；是在节省蜡的自然选择压力下的几个进化阶段形成的。