Emphasizes ethological studies of natural behavior patterns and their analysis in laboratory work, with contributions from field biology (mammology, primatology), sociobiology, and comparative psychology.

• 强调自然行为的动物行为学研究和实验室分析，以及田野生物学(哺乳动物学、灵长类动物学)、社会生物学和比较心理学的相关研究。