Eastern Atlantic and Mediterranean Sea: including the Adriatic, Aegean and Black Sea, Portugal and Senegal. Has also been reported from southern Morocco, Mauritania, and Angola (Ref. 10813).

• 东大西洋与地中海:包括亚得里亚海，爱琴海与黑海，葡萄牙与塞内加尔。已经也被记录自摩洛哥南部，茅利塔尼亚与安哥拉。(参考文献10813