Despite the growing evidence against a long-lived seaway in Miocene Amazonia, the strontium signatures in the mollusks revealed that this enormous lake system did experience occasional salinity increases.

• 尽管越来越多的证据，反对亚马逊河流域在中新世有长存的海道，但是软体动物壳上的锶标记，却还显示出这个庞大的湖泊系统，的确经历了几次盐份升高的过程。