Columella 1. The structure present in sporangia of many zygomycete fungi (e.g. Mucor) produced by formation of a domeshaped septum cutting off the sporangium from the sporangiohore.

• 1囊轴：许多接合菌的孢子囊中的结构（毛霉属）。在孢子囊的中央形成一个中空的腔，其外一层薄壁就叫做囊轴。