Abnormality of tapetum :The tapetum in JIYUGENG began to break up at meiosis stage, and basically disintegrated at late microspore stage.

• 2．药壁组织绒毡层结构的异常：可育品种吉玉粳绒毡层从减数分裂期开始降解，到小孢子晚期基本上解体；